Cephaloziella divaricata

Cephaloziella divaricata
Phylum: Marchantiophyta
Genus: Cephaloziella
Synonyms: Cephalozia aequitexta Steph. ex Bonner, Cephalozia asperifolia (Taylor) Trevis., Cephalozia asperifolia fo. belsensis Douin, Cephalozia divaricata (Sm.) Dumort., Cephalozia divaricata var. scabra M. Howe, Cephalozia notarisiana C. Massal., Cephalozia papillosa Douin, Cephaloziella aequitexta Steph. ex Fulford, Cephaloziella andina Herzog, Cephaloziella brasiliensis S.W. Arnell, Cephaloziella byssacea (Roth) Warnst., Cephaloziella byssacea var. asperifolia Macvicar, Cephaloziella carnutensis Douin, Cephaloziella divaricata var. asperifolia (Taylor) Damsh., Cephaloziella divaricata var. incurva (Lindb.) C.E.O. Jensen, Cephaloziella divaricata var. polystratosa (R.M. Schust. & Damsh.) Potemkin, Cephaloziella divaricata var. rupestris (C.E.O. Jensen ex Arnell) Damsh., Cephaloziella divaricata var. scabra (M. Howe) Haynes, Cephaloziella latzeliana Schiffn., Cephaloziella limprichtii var. stellulifera (Taylor ex Carrington & Pearson) Müll. Frib., Cephaloziella nigrimonasteriensis Douin, Cephaloziella papillosa (Douin) Schiffn., Cephaloziella rupestris (C.E.O. Jensen ex Arnell) Schumack. & Váňa, Cephaloziella starkei Schiffn., Cephaloziella starkei var. asperifolia Macvicar, Cephaloziella starkei var. examphigastriata Douin ex Müll. Frib., Cephaloziella starkei var. papillosa (Douin) Casares-Gil, Cephaloziella starkei var. scabra (M. Howe) L. Clark & Frye, Cephaloziella starkei var. serriflora Douin, Jungermannia asperifolia Taylor, Jungermannia byssacea Roth, Jungermannia confervoides Raddi, Jungermannia divaricata fo. laxa Nees, Jungermannia divaricata Sm., Jungermannia intricata Huebener, Jungermannia starkei fo. procerior Nees, Jungermannia starkei Funck ex Nees, Jungermannia stellulifera Taylor ex Carrington & Pearson
Common NameCommon threadwort
Quick & Dirty IDXXXXXX
© John Reynolds
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© Connor Wardrop

Distinguishing Features

Small enough to evade notice (<.5 mm wide) but ambient enough to be found in almost any habitat, C. divaricata varies from green to black in colour and solitary threads to dense tufts in growth habit. The vegetative leaves are smaller than the shoot diameter and are sharply two-lobed to about 1/2 or 3/4 leave length and fairly smooth along their edges. Microscopic examination should be used to confirm this latter character as well as the presence of oil bodies.

Similar species

Sphenolobopsis, Marsupella, Cephalozia and Fuscocephaloziopsis can be of similair size and leaf form, but the leaves in those genera are wider than the stems and have oil bodies, while in C. divaricata oil bodies are lacking and the leaves are narrower than the stem. Eremonotus myriocarpus is very similar in size, leaf shape and threadlike growth habit. It grows in the very specific habitat of damp vertical rock, but otherwise requires granular examination of the shoots to compare leaf shapes and reproductive structures- see photos under that species. Other species of Cephaloziella (C. phyllacantha, C. spinigera, C. turneri, C. elachista) in this region have leaves divided almost to their bases in addition to prominent teeth along the leaf margins. Vegetative leaves of C. divaricata are divided to about half their length and typically without teeth. The remaining species will need to be parsed based on the arrngement of their reproductive organs. In Cephaloziella divaricata, the modified leafy structures that bear the archegonia (“gynoecia”) and antheridia (“androecia”), while in C. rubellum, C. varians and C. hampeana, they can be found on the same shoots.

Habitat

Dry to wet humus, mineral soil, decayed wood, rock outcrops, boulders, cliffs, ledges, crevices, streambanks, waterfall spray zones, bogs, disturbed areas, and occasionally tree trunks in the lowland, steppe, montane, and subalpine zones; common in coastal BC, infrequent in sc BC, rare in se and n BC.

Associated species

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Distribution Map

Relevant Literature