Scapania undulata

Scapania undulata
Phylum: Marchantiophyta
Family: Scapaniaceae
Genus: Scapania
Synonyms: Jungermannia cochleariformis Weiss, Jungermannia nemorosa var. purpurascens Hook., Jungermannia purpurea Scop., Jungermannia resupinata L., Jungermannia undulata fo. purpurea Nees, Jungermannia undulata fo. speciosa Nees, Jungermannia undulata L., Martinellia dentata (Dumort.) Arnell, Martinellia nemorosa var. purpurea Lindb., Martinellia purpurascens (Hook.) Arnell & C.E.O. Jensen, Martinellia purpurascens var. ambigua (De Not.) C.E.O. Jensen, Martinellia undulata var. purpurea (Nees) Lindb., Martinellius resupinatus (L.) Gray, Martinellius undulatus (L.) Gray, Mnium jungermannia L., Plagiochila undulata (L.) Mont. & Nees, Pleurozia cochleariformis (Weiss) Dumort., Radula cochleariformis (Weiss) Dumort., Radula dentata Dumort., Radula undulata Dumort., Scapania alata Steph., Scapania atrata Warnst., Scapania dentata (Dumort.) Dumort., Scapania dentata var. ambigua (De Not.) C. Massal., Scapania dentata var. heterophylla Müll. Frib., Scapania dentata var. speciosa (Nees) Cogn., Scapania dentata var. taeniiformis Müll. Frib., Scapania douinii Schiffn., Scapania falcata Steph. ex Müll. Frib., Scapania gigantea Horik., Scapania groenlandica Steph., Scapania informis Steph., Scapania intermedia (Husn.) Lamy, Scapania intermedia (Husn.) Pearson, Scapania komagadakensis Amakawa, Scapania nemorosa var. intermedia Husn., Scapania nemorosa var. purpurascens (Hook.) Gottsche, Lindenb. & Nees, Scapania nuda Steph., Scapania oakesii Austin, Scapania oseensis Warnst., Scapania paludosa var. aquatica Schiffn., Scapania paludosa var. natans Schiffn., Scapania purpurascens (Hook.) Taylor ex Pearson, Scapania purpurea Kashyap & R.S. Chopra, Scapania resupinata (L.) Dumort., Scapania speciosa (Nees) Lett, Scapania splendens Steph., Scapania squarrosula Lindenb., Scapania suecica Warnst., Scapania undulata var. aequatiformis (De Not.) Müll. Frib., Scapania undulata var. alata Schiffn., Scapania undulata var. ambigua De Not., Scapania undulata var. densa Müll. Frib., Scapania undulata var. dentata (Dumort.) McArdle, Scapania undulata var. flagellaris Loeske, Scapania undulata var. integrifolia Lindb., Scapania undulata var. lepida Warnst., Scapania undulata var. oakesii (Austin) H. Buch, Scapania undulata var. purpurascens (Huebener) Cogn., Scapania undulata var. purpurea (Nees) Nees, Scapania undulata var. speciosa (Nees) Nees, Scapania undulata var. subdenticulata Warnst., Scapania undulata var. subglacialis Schiffn.
Common NameWater Earwort
Quick & Dirty IDSmooth Compressed Water Scapania
© John Reynolds
iNaturalist
© Randal Mindell
iNaturalist
© Julia Carr
iNaturalist

Distinguishing Features

Scapania undulata is the most common Scapania in habitats that are at least seasonally inundated with water. The flaccid shoots are little-branched and elongate (to 4 mm wide, 3+ cm long) and vary from green to red to purple and almost black, with the uneven and ellipsoidal leaf lobes becoming tightly appressed to one another near the shoot tip. Leaf lobes can be toothed or untoothed and the upper lobe varies from about 1/3rd to 2/3rds the length of the lower lobe, the two lobes converging along a sharp fold (“keel”) that is about 1/3 the length of the lower lobe.

Similar species

Scapania uliginosa and S. paludicola can be found in similair wet habitats, but both of those species have an arched keel, while the keel in S. undulata lacks the dramatic arch. Because it is a highly variable species in terms of color and the presence and absence of teeth, care should be taken to note the attachment of the upper and lower lobes to the stem. The upper lobe attaches directly across the stem (“non-decurrent”) while the lower lobe attachment runs down the length of the stem (“decurrent”). When teeth are present, these attachments help distinguish it from Scapania americana, which has an upper lobe that is decurrent. S. subalpina and S. obscura have similair lobe attachments and smooth margins, but their upper lobes are »3/4 the size of their lower lobes, whilst in S. undulata, the upper lobe is < 2/3rds the size of the lower lobe.

Habitat

Moist to wet mineral soil, rock outcrops, boulders, cliffs, streambanks, waterfall spray zones, bogs, and seepage sites, often submerged in shallow water, in the lowland, montane, and subalpine zones

Associated species

Pellia neesiana, Racomitrium aciculare, Marsupella emarginata

Distribution Map

Relevant Literature